As per profJim's questions really.
DO you have a PC Speaker plugged into the board's speaker port? Some cases have them either bolted to the case or they come in the accessories kit.
To be honest the only beep code I've ever gotten is the continuous short beeps. The manual says "Continuous short beeps: Power error" however there's nothing wrong with my power supply, connections etc, which means short beeps are also Morse code for "Aaaaagh! your settings are wrong, I'm not going to boot till you switch me off at the wall.... and then only from the backup BIOS"
I'd suspect RAM first off too. 5-4-4-12 is tight for a 100MHz (DDR) overclock. Your RAM could verry well be up to it, heck if it's D9 IC's then it'll do 5-5-5-15 @ 1100-1200 DDR. You also have 4 sticks. It may be possible that it wants another tick or two of MCH Core voltage.
Also if the BIOS info you posted is 100% as it apears in your bios, the board, if left to its own devices would set tRead=10. I'm not saying it's right, but it could be (sounds like 9 SHOULD be fine, 450 FSB is hardly huge).
Without altering any voltage settings, set your RAM to 5-5-5-15/18 and tRead to 10.
Can you boot now? If yes, then change each timing back one at a time, 5-5-5-12 then 5-5-4-12, then 5-4-4-12, tRead back to 9. Actually change tRead back first. 99% of my reboot loops and beeping fits have been through damned bloody stupid tRead setting.
To try and eliminate the processor as the culprit, you could lower your multiplier to 8. If it boots with your ambitious RAM timings then you could need more vCore, VTT etc.
Final suggestion: you have a FSB hole. It's not serious, no doctor required. Try raising FSB to 455 or 460, lowering CPU multi if you think you might have processor issues if jumping FSB speed up rapidly.
Any time you hit a wall in overclocking, look for the weakest link. Look over BIOS settings and think "What am I pushing my luck on? Have I tightened stuff up so much it's fit to burst or do I have something clocked past specs other than the CPU?".